#NEXUS BEGIN DATA; DIMENSIONS NTAX=39 NCHAR=323; FORMAT DATATYPE = STANDARD GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4 5 6"; MATRIX Teius_teyou 00000?0000000000010000001?01000000000010000001000010?000?01000000011010000000100000100100000000000000000??0110010100000102000110000000110000000000000000001000100100000??002100000000000310000012010011100121000100000000110001?11100?000000000101010000110020000000001110011001101100000000011111{1 2}00100001000001021000000010000000 Takydromus_sexlineatus 00000??0001020000102100000010000100000002200030?0000?001?00000000001000000000100000000100000000000000000??0010010100000000002100000000?00000000000000000001000000000000??00200000100000021000001111000110012000000000000?000001?00100?00000?00010000100001??1?0??01?001{0 1}1001100111110000{0 1}00?010011{0 1}0010000111000021000???????000000 Lacerta_viridis 00000??00000200001?1100000000000101000100200030?0000?001?000001010110000000000000000001000000000000000000?0010010100000000001100000000?00000000000000000001000000000000??002000001000000310000011010001100120000000000000000001000000?00000?000100001000010010000000001110111001111100000000010011100100003110000110000?0?01?000000 Scincoideus_grassator 00000?????0????????????002???0???????010????030???0?????0???0000101??0?????????000?0?????????????????????????????????????????????????????????????????????????????????10??0020??01???0???3?????????????????????????????????00000010000??0000?000???????????????????????????????????????????????????????????????????????????????????? Cryptolacerta_hassiaca 000?0????0?010?0?1????0002?0000010?0001?0210020???00?0?0?0?100????1?000?11000??000000010?????0?????????0?????????????????????????????????????????????????010?????????10????210?01100??0?2??????1??110??????0????10??????0?00001010000?20000?000??????00???00?00000???0???01???0??????000?00001?????0???0??211???????0?????????????? Jepsibaena_minor 1{1 2}00???{0 1}?02??1??????????0????????21??????????0??00?0??1??02?210??????0?12???????????????1?11???1???22?0????0????10100?11100110111??0??0?????011111?????11101?111???1?11????200?0011022?00???0001???000?????2??1?10101?0111??010131?11?2?1112000?????????????????????????????????????????????????????????????????????1?????????????? Dyticonastis_rensbergeri 120020?11021021?00030101010100111210102100111000001001100020220100110011220001?0011110??1?11???????2220????011011010011110111011100002011113011111101111110111111?1111110?01010002102210010000011110001100120110001011011011010131011?211112000???????????????????????????????????????????????????????????20??11????1?????????????? Spathorhynchus_natronicus 1200?????0?002?0?00???0?0??100111????021001?100100100110?02?220???11011122200?1001111???1??1?????1?2210?????????1?10??1110?1101?10?0???1?1?30111111????1?101?111???1?11?????00?00110???????????1??1000?1???2????101?110110??010131?11?22???????????????????????1??????????????????????????????????????????20????????1?????????????? Spathorhynchus_fossorium 12002{0 1}?1102102110003000101010011121010210011100100100110?020220?001101112220011001111011101100110?0222010100110110100111101110111000020111130111111011111??11111?11111110?01010?01102210000000010010001100120110101021011011010131011?221112000???????2?100100?11?????????????????????????????????????????20??11????1?????????????? Hyporhina_galbreathi 120011?1102202?1?00???0?0{0 1}?000?11?1??0221?1?100100110100?02?23010?11010112000?10011101??1??1??11?1?2220????0??0?10100?1?10?11?1?10??02?????30111111??1?1{1 2}??1?111???1????????????????????????????????????????????????????????0??141?20?2???????????????????????????????????????????????????????????????????0?????????1?????????????? Hyporhina_tertia 120011?1?02202?1?00???0?0??000??1????02?????????????????????23010?1101?1???00?1????????????????????????????011???010???1???1?????0??0??1???????????????????????????????????2????????????????????????????????????????????????010141?20?2???????????????????????????????????????????????????????????????????0?????????1?????????????? Hyporhina_antiqua 120011??1?2102?1?00???0?0??000111?1??0221?1?100100110100?02?23010?11010112000?1001110111101111?????22?0???00????1?100?11?0?11?1?10?0??01?1?3011?111????1{1 2}101?111???1?11?????????0?????????????????1??0?1???{1 2}????101??1??10??010141????231???0?0???????????????????????????????????????????????????????????0?????????1?????????????? Hadrorhineura_hibbardi 1100?????????2???0??????0????011121??0211?1?100?00100110?02??{0 1 2}0???11?00?2??00?1?1????1?11011?1??0??2220???0?????????????????????10???2?1?1?30101111??1?1???1?1???????1???????0?00110??????????????1000?1???{1 2}??1?101??1011???010????11?2311120?0???????????????????????????????????????????????????????????2?????????1?????????????? UCM_19852 1100???????012?0?0????0?0??10011121??0211?1?100100100?10?02??10???11001?21000?1?111101??1111???1?1?2220?????1?0???100?1??0?11???10?0???1?1?30111111????1{1 2}??1?111?????11?????0????1?????????????1??100??1???2??1?10???1??????01014??21?221?????????????????????????????????????????????????????????????????2???????????????????????? Protorhineura_hatcheri 110011?11020021000?3010?01010011121010211?1?100?00?00110?0202101001100112200011011110111101100010?02220???0011??101002111011101110000201111301111110111121011111???1?1110?020000011022110100000??01000110?12?100101011011011010131121?231112000???????????????????????????????????????????????????????????20??????????????????????? Rhineura_floridana 110011?1102002100003?10101010011121010211?1010010110011000202301001100111220011011111011101100010102220101001101101002111011101110000201110301111110111121011111111111110?011000011022110110000??010001100111?10101011011011010141?40?2411120000?????42210010011201111???2????1?????1????1?21??????????1??200111100?211001100110111 Blanosaurus_primeocaenus ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1?011211?1?1?10000?????????????????0??0??????????????1?010????????210??010????????????????01?????????????????????????????????????????????????????????????????? Blanus_strauchi 00000?0010221100?00?100102100000121?10301?1?001011101010012?00011011000011100?11????10111110001001010200???0100?11101??1?001111110110112001301111110?1110111?110???1?110112{0 1}001011110000110001010011010000?00210101000111011010110141113101{0 1}130????????1?????1?02???????????????????????????1?????????????000111????11???????100101 Blanus_mendezi 00000?00002211000002100?00100000121010301?10001011101010012?00011011100011000?11????1?11111??0?????????0???0?00??11???10??0?111110?101120?1301111110?1?10101?110???1?11011210010111100000?00???10?11010000100?101010001110110101101310121211130?????????????01?02?????????????????????????????????????????0???1?????1?????????????? Blanus_cinereus 00000??0?0221100?00??00?02?00000121?10301?1?001011101010012?1001?111000011100?11????11111110001001010200???0100?11101??1?001111110110112?0132111111??1?10101?110???1?1101120101021100000100001010011010000?00?0010100112101101?11014111210111300?011032???11?11020?1?????????????????0??21?11??????????0?1000111????110011011100101 Oligodontosaurus_wyomingensis ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????11?0??210?0111010000?0000000?10?0010?10??????????????11010????????1110?001???????????????????????????????????????????????????????????????????????????????????? Oligodontosaurus_sp ??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????1?1?0?1?1?1?0?0?????????????0????0??????????????11?10?????????1????01???????????????????????????????????????????????????????????????????????????????????? Anniealexandria_gansi 000?0?????2?????????????????????????????????????????????????101??2???????????????????????????????????????????????????????????????????????????????????????????????????110001210101???000?1?1???????????0?00?0?2?0??110?1???110101101?0??111??0?0???????????????????????????????????????????????????????????????????????????????????? Bipes_biporus 0000100001221100000010?11?010000131011??1?1?001011101011?12010111201000022?01111????10111110001001011200??0100111?1112100001211110010112011301011111111101001110?111111000111010111000011?11110?00110100001012101111001110010101{1 2}0140?1311110000011104211111011020112012011001000020110121110010002110101100011110?0010011011100100 Bipes_canaliculatus 00000?0001221100000010?11?010000131011??1?1?001011101011?12010111201000022?01111????10111110001001011200??010011101112100001211110110112011301011111111101001110?111111000211010111000011?11100?0011010000101210111100111001010110140?13111100000111032?11110?102011?0020110010000101101211100100021101011000111????010011011100100 Cadea_palirostrata 10110?0{0 1}11022110100?110?0{0 1}111200121??0301?1?001020101100112?0001??10000012200?11????11111111?011111??2211010101?11101?10?001011110111112011321111111?1?1030??111???1?110??1100111300200?0?0111?0??0111?00??0?2011110?1201011010111230?221011030???????2?????0????????1???2????1?????1???????1?????????????00?11?????1????????120101 Todrasaurus_gheerbrandti ?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????02?1??01???0?0????????????????????0??????????????10010????????4??????0???????????????????????????????????????????????????????????????????????????????????? Trogonophis_wiegmanni 00000?0111221100000010010?011001121010301?11001020101010112020010?1121002?21?111????10111111001101120201000000111011020011011111101?00120003010111111111010011101111111200210010230000011?01011???01110010001?1?111001111?10110120240?13110?1?00111?02?01111??1021?1?1???2????1?????111??1?21??????????1??0001111???110001011100101 Diplometopon_zarudnyii 1200{0 1}?00112212000000100100011011121010301?1010011010001111202001001121001??2??11????11111111110101120201010000101011010011011111100?00120113010111111111010011101111111200211010230000011?01010??001110010001?11?11101121?21112130141113110?100011???32010110?1001?1?003?1?0??1?????111??1?21??????????1??000111????110001011110101 Amphisbaena_fuliginosa 00000?10002111101010111100110000021010301?10001020101000012010111111101022101011????1101111100101112021110101111111000100001111110111212011311111111111102?01111?11111100022101123000001??111100?001111001101201?11001221?110101111310221011100?0????1?211110?1020?12?????????1?????1??????21??????????????00111100?110111011100101 Amphisbaena_alba 00000?00102211101010110102010000121010301?10001020101000012000011111001021101011????1101111101101112022110101111111001100001111110111112011311011111111102001111???1?11000221010230000010?111100?00111100110120111100122101101011113112110111000???????2??11??1020?1???????????????????????2????????????????01?0????1101110111?0101 Geocalamus_acutus 10110?1010221210100?100102100101121110301?1010001010000011201000021000001111??11????1101111111100112021110100011111012101001311110111112011321011111111103??1111?11111100020101113000001??1111100?01110000101201111001121?110101{2 3}12411131011100001110?????11??1020?1?1???1000?1?????1????1?21??????????1??000111????111111011120111 Loveridgea_ionidesii 10010?0010221200100010011?100100121010301?10100010101000112?100?0?10000011101011????11011111111001?202111010???1111???10??0??1?110?111?20113210111???1?10???1?11???1?11000211011230000?10?11?110??0111?000101201111001221011010?31?4??231???1?00???????2??11??1020?1???????????????????????2??????????????000111????110111011120101 Amphisbaena_kingii 10110?1011022210100011010?101200121110301?10001020101000012?10000?10100011201111????11011111111101120221101010111110121010010111101112120113211111111111030?1111???1?11000211011230000010?111110??01111000?012011110012210110101112411231011100000???32???11??10?0?1???????????????????????21??????????1??00?111????111111011120101 Lophocranion_rusingense ???????????????????????10????0?0?2??10301?10000020101000012???????????????{0 1}?????????11??1??1?????112021????????????????????????????????????30111111111110{2 3}0?11?????1??????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????0????1??????????????????? Leposternon_microcephalum 11000?0210022210100010010{0 1}100001121010301?10001020100000002?12010?11201112201111????1111111111110112020111101?11101?1?11?0011111111111121103211111111111030?1111???1?1?00021??112300000100110110??0111?000100201?1?001121??10???2114112410101000??????????11??1120?1???????????????????????2??????????????0??1?1????111001000110111 Listromycter_leakeyi 11000?01100221001000100002110011121010301?1010002?1?0?0?????10010?11?001???{1 2}??11???????????????????????????0111110100211000?11???1?11202??1??1????1111?10???????????????????????????????????????????????????????????????????01?11123??2?????1?????????????????????????????????????????????????????????????2???1?????1?????????????? Monopeltis_capensis 11000??2100022???00???0?0??000111?1??00?1?1?0010?0100000002?23010?110?112?110?11????10111011101001?2020???001?1?10110?11?0011111111?1?12?1130111111??1?12301?111???1?1?????2???12??0?00???11??????????????????????1??11????1010141?410231???100??????32???11??1120?1???????????????????????2??????????????0?????????211011000111111 Monopeltis_jugularis 11000??2100222????0???0?0??00?111????00?1?1?0010?0100000?02?13010?110?111221???????????????????????????????????????0??1??????????????????????1?????????????????????????????????????????????????????????????0??0?111??112???101001123?1231????00???????????????????????????????????????????????????????????0?????????1?????????????? ; END; begin mrbayes; constraint trogonophiidae partial = Trogonophis_wiegmanni Diplometopon_zarudnyii : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint amphisbaenidae partial = Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii ; constraint amphisbaena_higher partial = Amphisbaena_fuliginosa Amphisbaena_alba : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint amphisbaena partial = Amphisbaena_fuliginosa Amphisbaena_alba Amphisbaena_kingii : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Geocalamus_acutus Loveridgea_ionidesii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint amphisbaenidae_sam partial = Amphisbaena_fuliginosa Amphisbaena_alba Amphisbaena_kingii Leposternon_microcephalum : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Geocalamus_acutus Loveridgea_ionidesii Monopeltis_capensis Monopeltis_jugularis ; constraint amphisbaenidae_afr partial = Geocalamus_acutus Monopeltis_capensis : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Loveridgea_ionidesii Amphisbaena_fuliginosa Amphisbaena_alba Amphisbaena_kingii Leposternon_microcephalum ; constraint amphisbaenoidea partial = Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata ; constraint bipes partial = Bipes_biporus Bipes_canaliculatus : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint blanus partial = Blanus_strauchi Blanus_cinereus : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint bipedoidea partial = Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint amphisbaenomorpha partial = Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis : Teius_teyou Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana ; constraint amphisbaenia partial = Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis : Teius_teyou Takydromus_sexlineatus Lacerta_viridis ; constraint lacertidae partial = Takydromus_sexlineatus Lacerta_viridis : Teius_teyou Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis ; constraint lacertibaenia partial = Takydromus_sexlineatus Lacerta_viridis Rhineura_floridana Blanus_strauchi Blanus_cinereus Bipes_biporus Bipes_canaliculatus Cadea_palirostrata Trogonophis_wiegmanni Diplometopon_zarudnyii Amphisbaena_fuliginosa Amphisbaena_alba Geocalamus_acutus Loveridgea_ionidesii Amphisbaena_kingii Leposternon_microcephalum Monopeltis_capensis Monopeltis_jugularis : Teius_teyou ; calibrate Scincoideus_grassator = Unif(52.5, 54.0) Cryptolacerta_hassiaca = Unif(47.4, 47.8) [general Messel, unknown level] Jepsibaena_minor = Unif(51, 52.5) [Lostcabinian, following Smith et al. 2008] Dyticonastis_rensbergeri = Unif(18.2, 26.94) Spathorhynchus_natronicus = Unif(35.57, 35.97) [between Ashes B and G, by Ar/Ar from Swisher & Prothero 1992] Spathorhynchus_fossorium = Unif(50.61, 51) Hyporhina_galbreathi = Unif(33.3, 33.7) [Orellan fide Taylor] Hyporhina_tertia = Unif(33.7, 37) [Chadronian age of holotype] Hyporhina_antiqua = Unif(30.8, 33.3) [Whitneyan fide Taylor 1951, PBDB] Hadrorhineura_hibbardi = Unif(33.3, 33.7) [Orellan fide Taylor 1951] UCM_19852 = Unif(33.3, 33.7) [Orellan fide Longrich et al. 2015] Protorhineura_hatcheri = Unif(30.8, 33.7) [Orellan-Whitneyan fide Taylor] Blanosaurus_primeocaenus = Unif(47.8, 56.0) [Dormaal MP7, Conde-en-Brie MP 8-9, Premontre MP 10; i.e., Ypresian, Auge 2012] Blanus_mendezi = Fixed(11.6) [Bolet et al. 2014] Oligodontosaurus_wyomingensis = Unif(57, 58.5) [Princeton Quarry, Ti-5a in C25r; check new Gradstein et al.] Oligodontosaurus_sp = Unif(61.7, 63.3) [Medicine Rocks fauna, Torrejonian, PBDB] Anniealexandria_gansi = Unif(55.8, 56) Todrasaurus_gheerbrandti = Unif(56, 59.2) [Thanetian fide Auge & Rage, 2006] Lophocranion_rusingense = Unif(17, 20) [Peppe et al. 2017] Listromycter_leakeyi = Unif(17, 20) [Peppe et al. 2017] ; outgroup Teius_teyou; ctype ord: 2 5 11-14 30 34 39 42 46 49 61 62 66 73-76 100 103 118 122 125 134 136 153 154 171 172 177 178 181 182 185 204 206 213 215 216 225 228 231 232 234 246-248 253 257 261 264 266 281 284 291 299 306 307 309; set autoclose=yes nowarn=yes; lset nst=6 rates=gamma; prset clockvarpr = igr; prset clockratepr = normal(0.0025, 0.1); prset topologypr = constraints (trogonophiidae, amphisbaenidae, amphisbaenoidea, amphisbaenidae_afr, amphisbaenidae_sam, amphisbaena, amphisbaena_higher, bipes, blanus, bipedoidea, amphisbaenomorpha, amphisbaenia, lacertidae, lacertibaenia) ; prset brlenspr = clock:fossilization; prset nodeagepr = calibrated; prset samplestrat = diversity; prset sampleprob = 0.018; [= 18/ c. 922 species] prset speciationpr = uniform(0,10); prset extinctionpr = beta(1,1); prset fossilizationpr = beta(1,1); prset treeagepr = offsetexp(97,180); [boreoteioids, Zheng-Wiens] mcmcp ngen= 20000000 relburnin=yes burninfrac=0.25 printfreq=5000 samplefreq=5000 nchains=4 savebrlens=yes; mcmc; sump; sumt; end;