#NEXUS BEGIN DATA; DIMENSIONS NTAX=54 NCHAR=175; FORMAT DATATYPE = STANDARD GAP = - MISSING = ? SYMBOLS = " 0 1 2 3 4"; MATRIX Polychrus_acutirostris 021200?0111001111110000000000011100000000011001011101?00001101001111010021010000?11110101110000210001??01110010101022?1?000000101212021221?110011110000000100?00100000010200000 Parasauromalus_olseni ?10000?0?01001001110?????000?111100010000011?010?1101000002001001111010021010000?0???0001????????????????0?0020101011?1?000010?01?????????????????????????????????????????????? Parasauromalus_LevelO ?????????????????????????????????????????????????????????????????????????????????????????????????????????????20101011?????????????????????????????????????????????????????????? Sauropithecoides_charisticus (1 2)012001011100111????00000???1?1?1?00000????01010????10000???010011010100210110????????????????????????????????00?1022????00?????1?????????????????????????????????????????????? Pristidactylus_torquatus 111000110000001000?000000001(0 1)001100000000000011020101000001000001101010(0 1)21000000?00100000??????????????000100101000211111000101012010010000010100100101100001000010101010200000 Anolbanolis_banalis 1100???0?0?00002?11?0000?000?02?1?10?00???01???0?010?0?00???100?11110000211000???????????????????????????????1010(0 1)011?????????0???????????????????????????????????????????????? Anolbanolis_LevelO ?????????????????????????000?02?1?????????????????????????????????????0?210000????????????????????????????????0???01??????????????????????????????????????????????????????????? Paranolis_LevelO ?????????????????????????0???0011??????????????????(0 1)?010??10????111000??(0 1)12000????????????????????????????????0???02??????????????????????????????????????????????????????????? Tuberculacerta_pearsoni 3(0 1)12001010100012?11?00000???001?1?00?00??????????0?010??1?11????111101???????????????????????????????????????00100000??1??????????????????????????????????????????????????????? Anolis_ricordi 011100101000001211100000000010011001?000000110101010101000??01001111010011210000?01110120??????????????010101111000220111110?0011?120111102110010100001100100?00110101010200000 Anolis_chlorocyanus 01000010?0000002111000000000(0 1)0111011?000000100101010101000??10001111010011000000?10000120??????????????010101111000220111100??011?120010102110001000000100100?001101010102000?? Anolis_cristatellus 01000000?0000002111000000000(0 1)0011001?00000010?1020101010000?10001111000011200000?10000120?10000211011??010101111000220111100??0112120011102?10010100000100100100110101010200001 Basiliscus_galeritus 021100100000001000?0001100000011100110000100111010111?10001111001111000031200100?00110100??????????????00000020100011110000010101??2021000?101000000001100011000010010010001011 Basiliscus_basiliscus 011100100000001010?00011000000111001100001001110111?1010001111001111000031200100?00110110100002100000??000000201000211100000??1?120?02200001010001000??100011000010010010001011 Oreithyia_oaklandi (0 1)11(0 1)011000000010?1??00(0 1)1?000111?1?1?2000???0?111?0??1111??11110?111111???1210????????????????????????????????10100001?10?00?????1?????????????????????????????????????????????? Corytophanes_cristatus (0 1)2110?101000001001000011000001111001200000011?1011111112002102111111000031210110?11110100100002110000??101001201?002210000001?001212021011?101010?00?01100000000010000010001011 Laemanctus_longipes (0 1)1010010000000100110001100000001101120000100111010101112001101001111110(0 1)(2 3)1210100?11110100??????????????100000101000211100000?0101?02021?11?101001?00001100000000?10000010001011 Geiseltaliellus_maarius 11100??0?000001010??00??00001011100000?0?100?010?01110(0 1)0?010010?111100002?2000?0?00?00100???????????????0??0?10100000?1?01100?101??2??10?1?10100010000?1??????0??????????????11 Suzanniwana_patriciana 1?10?????0000010????00(0 1)0?00010111?002000?1?01?10?010?0000?10?10?111(0 1)0100212000??????????0????????????????????101000001???000??1?1?????????????????????????????????????????????? Suzanniwana_LevelO ?(0 1)?????00010001(0 1)?0?000(0 1)0??00?0111????????????????0?(0 1)?00???10???01111010?2???????????????0????????????????????10100000?100???????1?????????????????????????????????????????????? Crotaphytus_collaris 11000?000000001200?001110100000010000000000001101?121000011000001100010020000000100000100210001110100210??00011101011110001010101001011001?000000?01101100000100000000010000011 Gambelia_wislizenii 1000001000000012010001010100000010002000000001101?121000011000001100000020000000100100000??????????????00000010100011110000010011?010110000101000101101100000100000000010000011 Cypressaurus_sp ??0000000000001100?00001110010001000?00?0?00?110{0 1}?1210001110??0011000000210000001000??1?0???????????????000001010?011010001010001?????????????????????????????????????????????? Queironius_praelapsus 1(1 2)00110000100011?0??00011101101?1?01201??0?0???0?010100?1?10????11(0 1)0000?200000???????????????????????????????101000220?0?10?????1?????????????????????????????????????????????? Dipsosaurus_dorsalis 130111000000001000?0000111001011110120100000000020101000111012001100000030000001000000110????????????2100000020210022010010000001?0201100010?0100001001100011000000000000000110 Brachylophus_fasciatus 131010?00000001000?000?101010011110110000000?00000101000101112001100(0 1)000200000010000001102100010101001100000020210022?1?01000010111202100110100?0??10??100000000000000000000110 Iguana_iguana 131100?00000001010?000010101(0 1)011111000000000000010101?00101111001100000021200001000100110110000010100110??00020210022010010010101?1202100110100001011011000000000000000000001?? Hoplocercus_spinosus 211100110000001100?0001?00000011?????0?000000?0?10001?0?10110??11100?0001000?000?00001100???????????????01???00100000010010110101??????????????????????1000???11???00????????00 Morunasaurus_groi 01110?100000001100?000001000(0 1)011?????0?010000???10101?0?10??0??11100?0002000?000?00001100???????????????0100020110000110010110111??????????????????????1000???11????0?????????? Enyalioides_oshaughnessyi 01110??10100011100?000?000001011100110001000?01010101000101101011110110020000000?00011100??????????????00100020(1 2)10000110010110101??10??200?1????0?0????100000010000000010000000 Oplurus_quadrimaculatus ?11?0??000?1001?0??001???000?010?01??0?00000????2?1?1?1?100?0?001100?0002?00?000?00010000110001110101210??00010100011???10000?1110020?1?0???20000?01011101011101010001110000000 Oplurus_cuvieri 210100000010001100?001010100101010010000000000101?121000100?11001100010010000000?00000110??????????????0??00010100000101100001111?0200100001200001010111010111010100011100000?? Chalarodon_madagascariensis 21010??00000001100?001?111000010101000000100?0102?121000101011001101000020000000?00000100??????????????0??000101000211110000011110010011000120001101000101011000010001110000000 Stenocercus_scapularis 1?000????00?000?0??001???00??010?0?????00000????20101?0?00100??01111?010??00?000?00000100??????????????00000010100022?1?100001011?010?1101?1?010110111?1?001100?1100000001000?? Microlophus_occipitalis 21000?10?0000012?1000100010000101?1000000000001020101000100?100?110100001000000??00000100??????????????001000101000220101000?1011?0100110000001001011??1?00110001100000?0200000 Tropidurus_torquatus 31000?10?0100012?10001000100(0 1)0101?1000000?00001020101000100???00110000001000000??00000100??????????????00000020100022010100???0?1?0100120000000000?11??1100110001100000002000?? Plica_umbra 21110?00?011000201000100110000101?1000000?00001020101000100?0101110000001000000??00000101????????????10000000101000220111000?1111?01001?0000010000011??1100???00110000000200000 Phymaturus_palluma 3301000000000011?0?00000110000101?10000??0?000102???10001?0?100?1100000022000000?1000?1002000011101002100100020100000010010???0?120000000000000000100??1000??100111000010000000 Liolaemus_pictus 2100110000000010?0?00100010000101?10000????000202??010101?0?110?1100000022000000?0000?100??????????????01000010100022011110???0?1?0?001?0000001011001??100011100111000010000000 Leiocephalus_melanochlorus 110010000010001100?0010101000010101000000100111020101010100?100011110101(2 3)010000??00000101??????????????000000201000220(0 1)0110001111?11001100000010110101?1000110001100000000000?? Leiocephalus_personatus 210010000010001100?0000101000010100000000000001020101000100?110011000000(2 3)0000000?0000010120000101010021000000201000220100100010111110011000?0010110101?1000110001100000000000?? Phrynosoma_platyrhinos 2001001000?00011010011000000011010?1000100???00022101001102111111101010022011000?100101?10?0010210100220??00000000000011100001011201001111?000000010010100011110000000001010000 Petrosaurus_thalassinus 11000???00100011011001?001001010101100000000?01020101001100?00001100000?20010000?000001011100102101002200100010100011?1?0000000111000010000000000000010110011100000000001010000 Sceloporus_undulatus 21000?100000001201000100000010101011000?00?000102?1?1000100?11001100000??0000000?00000121010??0210000220??00010100011011000???1?1?01001100000010010101?110011100100000011010000 Physignathus_cocincinus 4?00001100011000010100010010000000010100100000001?121000010?010011110000?0000000?0011010100110100000000000110?10?0100010100110011202002121?101010000020100000000000000010001010 Agama_agama 4?010001000110000101010100110000100101001000?0001?121000010?01001110000120000000?0010010101110010000?220?0110?100010000110011?0012?2001121?1?001001002010?0??100????0????????00 Priscagama_gobiensis 3?000??000?010?0010101??010000001001?0000000????11?010000110000?111001002?010000?001?0100???????????????0??00?100?100?1?0000?01?1????????????????????????0????????????????????? Leiolepis_belliana 3100001100011?0000?1110101001010000100100000?001??1210000110110011(0 1)00011?0000001?001?1101????????????10010000?110010000?100110001?02001101??0100110002?100000100000000010011000 Uromastyx_acanthinura 4?0100000000100100?11101101011111001000100?0?00010111000?10?010011000011?0011000?101011?12111010001001100?00??10?01000001001100?100200110(0 1)?0000011010(0 2)0100000101000000010001100 Brookesia_superciliaris 4200100?10?01100010111010000?011000001?110?1??202????000002?0110111101003?01100????1??1?1??????????????10??00?11001000011001100002??101211?01??????00??000000000110?0?110001000 Tinosaurus_LevelO ??????????????????????????????????????????????????????????????????????????????????????????????????????????????1100100?????????????????????????????????????????????????????????? Tinosaurus_MPH 30?(0 1)???????????????11?0??10???????0???1???????????????????????????????????????????????????????????????????????11001???????????????????????????????????????????????????????????? Saichangurvel_davidsoni 0(0 1)000??000000?0?0????????100?010?00??0?00?00????00000?0??0??1?0?110??00?1?00000????????????????????????????00101??0???1?????????1?????1?0000????????0?????????????????????????0 Elgaria_multicarinata 010000000010000000?00000?00000010011?00001?0012000000000?01021000001010001011000?00000100????121????1000000001001100000101000000??0?00110020?01011000000000??1000?0?00?0??0??10 ; END; begin mrbayes; constraint leiocephalus partial = Leiocephalus_melanochlorus Leiocephalus_personatus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint tropidurinae partial = Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint oplurus partial = Oplurus_quadrimaculatus Oplurus_cuvieri : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint oplurinae partial = Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint leioplurinae partial = Pristidactylus_torquatus Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis : Polychrus_acutirostris Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint dipsolophus partial = Dipsosaurus_dorsalis Brachylophus_fasciatus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint iguaninae partial = Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint hoplocercinae partial = Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint crotaphytinae partial = Crotaphytus_collaris Gambelia_wislizenii : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint laemophanes partial = Corytophanes_cristatus Laemanctus_longipes : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint basiliscus partial = Basiliscus_galeritus Basiliscus_basiliscus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint corytophaninae partial = Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint anolis partial = Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus : Polychrus_acutirostris Pristidactylus_torquatus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint polychrotinae partial = Polychrus_acutirostris Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus : Pristidactylus_torquatus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint higheriguanidae2 partial = Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Leiocephalus_melanochlorus Leiocephalus_personatus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint higheriguanidae1 partial = Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus : Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint petroporus partial = Petrosaurus_thalassinus Sceloporus_undulatus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint phrynosomatinae partial = Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint iguanidae partial = Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus : Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint agaminae partial = Physignathus_cocincinus Agama_agama : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint leiagaminae partial = Physignathus_cocincinus Agama_agama Leiolepis_belliana : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Uromastyx_acanthinura Brookesia_superciliaris Elgaria_multicarinata ; constraint agamidae partial = Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Brookesia_superciliaris Elgaria_multicarinata ; constraint acrodonta partial = Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris : Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Elgaria_multicarinata ; constraint iguania partial = Polychrus_acutirostris Pristidactylus_torquatus Anolis_ricordi Anolis_chlorocyanus Anolis_cristatellus Basiliscus_galeritus Basiliscus_basiliscus Corytophanes_cristatus Laemanctus_longipes Crotaphytus_collaris Gambelia_wislizenii Dipsosaurus_dorsalis Brachylophus_fasciatus Iguana_iguana Hoplocercus_spinosus Morunasaurus_groi Enyalioides_oshaughnessyi Oplurus_quadrimaculatus Oplurus_cuvieri Chalarodon_madagascariensis Stenocercus_scapularis Microlophus_occipitalis Tropidurus_torquatus Plica_umbra Phymaturus_palluma Liolaemus_pictus Leiocephalus_melanochlorus Leiocephalus_personatus Phrynosoma_platyrhinos Petrosaurus_thalassinus Sceloporus_undulatus Physignathus_cocincinus Agama_agama Leiolepis_belliana Uromastyx_acanthinura Brookesia_superciliaris : Elgaria_multicarinata ; calibrate Parasauromalus_olseni = Unif(52.5, 53) Parasauromalus_LevelO = Unif(53, 53.5) Sauropithecoides_charisticus = Unif(35.2, 35.7) Anolbanolis_banalis = Unif(55.8, 56.0) Anolbanolis_LevelO = Unif(52.5, 53.5) Paranolis_LevelO = Unif(52.5, 53.5) Tuberculacerta_pearsoni = Unif(35.2, 35.7) Oreithyia_oaklandi = Unif(35.2,35.7) Geiseltaliellus_maarius = Unif(47.4, 47.8) [marker beds M to Alpha] Suzanniwana_patriciana = Unif(55.8, 56.0) Suzanniwana_LevelO = Unif(53, 54) Cypressaurus_sp = Unif(31.034, 35.7) Queironius_praelapsus = Unif(35.2, 35.7) Priscagama_gobiensis = Unif(66.0, 83.6) Tinosaurus_LevelO = Unif(53, 54) Tinosaurus_MPH = Unif(35.2,35.7) Saichangurvel_davidsoni = Unif(72.1, 83.6) ; outgroup Elgaria_multicarinata; ctype ord: 1 4 16 37 49 52 56 59 62 73 75 90 95 96 102 110 112 117 121 132 134-136 170; set autoclose=yes nowarn=yes; lset nst=1 rates=gamma; prset clockvarpr = tk02; prset tk02varpr = exp(10); [default MrBayes] prset clockratepr = lognorm(-6, 1); prset topologypr = constraints (leiocephalus, tropidurinae, oplurus, oplurinae, leioplurinae, dipsolophus, iguaninae, hoplocercinae, crotaphytinae, laemophanes, basiliscus, corytophaninae, anolis, polychrotinae, higheriguanidae2, higheriguanidae1, petroporus, phrynosomatinae, iguanidae, agaminae, leiagaminae, agamidae, acrodonta, iguania) ; prset brlenspr = clock:fossilization; prset nodeagepr = calibrated; prset samplestrat = diversity; prset sampleprob = 0.02; [= 36/ c. 1975 species] prset speciationpr = uniform(0,10); prset extinctionpr = beta(1,1); prset fossilizationpr = beta(1,1); prset treeagepr = offsetexp(75,168); [ Flaming Cliffs, Zheng-Wiens ] mcmcp ngen= 20000000 relburnin=yes burninfrac=0.25 printfreq=5000 samplefreq=5000 nchains=4 savebrlens=yes; mcmc; sump; sumt; end;